Disentangling the effects of fishing and environmental forcing on demographic variation in an exploited species

a Department of Ecology, Evolution, and Marine Biology, University of California, Santa Barbara, CA 93106, USA b Tijuana River National Estuarine Research Reserve, Imperial Beach, CA 91932, USA c Facultad de Ciencias, Universidad Autónoma de Baja California, Carretera Transpeninsular Ensenada – Tijuana, No. 3917, Colonia Playitas, C.P. 22860 Ensenada, Baja California, Mexico d Leigh Marine Laboratory, Institute of Marine Science, University of Auckland, Auckland, New Zealand e Earth Research Institute, University of California, Santa Barbara, CA 93106, USA f National Center for Ecological Analysis and Synthesis, University of California, Santa Barbara, CA 93101, USA g Interdepartmental Graduate Program in Marine Science, University of California, Santa Barbara, CA 93106, USA h Marine Science Institute, University of California, Santa Barbara, CA 93106, USA i Moss Landing Marine Laboratories, Moss Landing, CA, 95039, USA j Bren School of Environmental Science & Management, University of California, Santa Barbara, CA 93106, USA